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According to this scenario, scales became feathers by first elongating, then growing fringed edges, and finally producing hooked and grooved barbules. 根據(jù)傳統(tǒng)理論，鱗片變成羽毛的第一步是先拉長，接著長出流蘇般的邊緣，最后產(chǎn)生有小鉤與溝槽的羽小枝。